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Lord Willas Tyrell is the eldest son and heir of Lord Mace Tyrell and Lady Alerie Hightower. When he was ten-years-old, his family arranged for him to be fostered with House Hornwood of the North after arranging a betrothal for both him and his sister, Margaery, in the North.

At nineteen, Willas was married to Lady Wynafryd Manderly and returned permanently to Highgarden to learn about ruling his family lands.

Unlike most of his family, Willas, who is the closest to his grandmother, Lady Olenna Tyrell, is aware of the plots to restore House Targaryen to the Iron Throne.

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