Ser Garlan Tyrell is the second son of Lord Mace Tyrell and Lady Alerie Tyrell. When he was young, he was named Garlan the Gallant by his brother, Willas, after he competed in a tourney at age nine. After Robert's Rebellion, Garlan was taken as a hostage by House Tully to keep the Tyrells in line, but he was released when he was fifteen for their good behavior.

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